THE SUBFAMILY DICHOMERIDINAE (LEPIDOPTERA, GELECHIIDAE): THE PHYLOGENY, CLASSIFICATION AND POSITION IN THE SYSTEM OF THE GELECHIIDAE

Ponomarenko M.G.

Entomologicheskoe obozrenie 2006; 85(2): 275-384

On the base of comparative morphological analysis taking into account the male genital musculature, the cladogram of the tribes and subfamilies of gelec-hiid moths is proposed. In accordance with the main skeleton branches of the cladogram the new system of the family Gelechiidae consisting of 5 subfamilies (Physoptilinae, Anomologinae, Gelechiinae, Anacampsinae and Dichomeridinae) is developed. The subfamily Physoptilinae is considered as a sister-group to other gelechiids, the monophyly of the latter is supported by the complex of synapomorphies (female retinaculum with anteriorly directed row of scales located on the base of radial stalk; muscles ma running across the longitudinal axis of the body, connecting the lateral arms of vinculum with margins of jux-ta; protractors of aedeagus /ra5 divided into two bunches mb& and m5b). The subfamilies Anacampsinae and Dichomeridinae are regarded as sistergroups. The monophyly of the subfamily Dichomeridinae is based on complex of synapo-morhies (presence of parategminal sclerites, which are the apodemes for muscles m4; presence of distinct ventral wall in tegumen; intrategminal position of the muscles m2). The subfamily Dichomeridinae is considered consisting of three tribes, Anarsiini, Chelariini and Dichomeridini, with 34 genera in total. The cladogram for the genera of the subfamily Dichomeridinae is proposed.