D. S. Aristov
Classification of the order Eoblattida (Insecta: Blattidea) with description of new taxa.
Number: 301, Pages: 1 - 56
The order Eoblattida (= Grylloblattida), assigned to the superorder Blattidea (Insecta), is revised. A total of 17 families are included in this order: the extinct Eoblattidae, Protophasmatidae, Cacurgidae, Mesorthopteridae, Ideliidae, Permotermopsidae, Euryptilonidae, Atactophlebiidae, Bardapteridae, Megakhosaridae, Blattogryllidae, Daldubidae, Idelinellidae, Permopectinidae, Soyanopteridae, and Doubraviidae and the extant Grylloblattidae. Fossil representatives of the order are known from the Paleozoic of North and South America, Eurasia and Africa and from the Mesozoic of Eurasia, Africa and Australia. Recent representatives of the order live in Canada, United States, Russia, China, North and South Korea and Japan. The family Taiophlebiidae from the Permian of Brazil is synonymized under Protophasmatidae, Permulidae and Kortshakoliidae from the Permian of Europe are synonymized under Permotermopsidae. Heterologellus (?) multidiffusus Brauckmann et Nerd, 2005 from the Carboniferous of Europe is transferred to the genus Protophasma (Protophasmatidae). The following new taxa of Eoblattida are described: Lomovatka udovichenkoi gen. et sp. n. (Eoblattidae; Carboniferous of Ukraine); Pintopinna gen. n. (type species Paranarkemina martinsnetoi Pinto, 1999, comb. n.; Protophasmatidae, Permian of Brazil), Suksunus bicodex gen. et sp. n. (Cacurgidae; Permian of Russia); Taskanatus gen. n. ( type species Alicula asiatica Storozhenko, 1997; Mesorthopteridae; Permian of Kazakhstan), Issadophlebia fusa gen. et sp. n. (Atactophlebiidae; Permian of Russia); Batkentak intactus gen. et sp. n. (Daldubidae; Triassic of Kyrgyzstan). Vologdoptera maculosa Aristov, 2009 (originally described in Kortshakoliidae) from the Permian of Russia is transferred to Orthoptera incertae sedis. The new Permian family Doubraviidae fam. n. is described to include Doubravia annosa Kukalová, 1964 (Sakmarian of the Czech Republic), Koshelevka gen. n. (type species Cerasopterum megakhosaroides Aristov, 2004; Kungurian of Russia); Iva permiana gen. et sp. n. and Belebey mutilus gen. et sp. n. (Kazanian of Russia). The origin of Eoblattida and its position within the infraclass Gryllones are considered; some aspects of the early evolution of Gryllones are discussed.