Diverse new Middle Jurassic Osmylopsychopidae (Neuroptera) from China shed light on the classification of psychopsoids

Peng Y.Y., Makarkin, V.N., Ren D.

В журнале Journal of Systematic Palaeontology

Год: 2016 Том: 14 Номер: 4 Страницы: 262-295

A well-preserved diverse fauna of fossil neuropterans of the family Osmylopsychopidae has been discovered at the Middle Jurassic locality of Daohugou, China. Daopsychops dissectus gen. et sp. nov., D. clausus gen. et sp. nov., D. inanis gen. et sp. nov., D. bifasciatus gen. et sp. nov., D. cubitalis gen. et sp. nov., Eupypsychops ferox gen. et sp. nov., E. confinis gen. et sp. nov., Nematopsychops unicus gen. et sp. nov., Ochropsychops multus gen. et sp. nov. and Stenopteropsychops trifasciatus gen. et sp. nov. are described. These new taxa allow a better understanding of the relationships between the psychopsoid families Osmylopsychopidae, Psychopsidae and Brongniartiellidae. The type of the latter (Brongniartiella gigas from the Late Jurassic of Germany) is redescribed. The venation of the new genera of Osmylopsychopidae shows that Brongniartiellidae is a synonym of this family. Of the genera ascribed to Brongniartiellidae, only Pseudopsychopsis belongs to Osmylopsychopidae with certainty. Gigantopsychops reticulatus gen. et sp. nov. from Daohugou (Psychopsidae) is an important taxon providing a new insight into the definition of the family. Wings of Psychopsidae may be distinguished from those of Osmylopsychopidae by a combination of the following character states: in both wings, the costal space is very broad distally; Rs1 is not deeply forked; the distal nygma is lost; the distal R1 is straight, not curved posteriad; in the hind wing, 1r-m is short and straight. It is clear now that the family Osmylopsychopidae was a dominant psychopsoid group during the Mesozoic, and Psychopsidae was relatively rare.

DOI 10.1080/14772019.2015.1042080

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